Proceedings of the National Academy of Sciences of the United States of America

About the PNAS Member Editor
Name Klein, Richard G.
Location Stanford University
Primary Field Evolutionary Biology
Secondary Field Anthropology
 Election Citation
Klein's research uses artifactual and faunal evidence to make significant contributions to our understanding of the evolution of human behavior, particularly the roots of modern behavior. His research documents important advances in hunting and gathering that enabled physically modern Africans to expand into Eurasia and replace the Neanderthals.
 Research Interests
I am a paleoanthropologist focusing on the fossil and archeological evidence for human evolution and particularly on the origins of fully modern humans. I have conducted field work in Spain and especially in South Africa, where I've analyzed animal remains from more than 100 archaeological and fossil sites dating between 500,000 years ago and the time of historic contact, about 500 years ago. I've shown how both species composition and average adult size in key mammalian species can be used to detect environmental change, including glacial/interglacial alternation, and I've developed criteria for distinguishing bone assemblages accumulated by people from ones accumulated by hyenas and other carnivores. However, I've worked mainly to illuminate long-term change in prehistoric human ability to hunt and gather. My research in South Africa implies that an especially important hunting-gathering advance occurred about 60,000-50,000 years ago. It promoted significantly larger populations and it has global implications, for it helps to explain how and why modern Africans expanded to Eurasia about the same time and how they then quickly replaced the Neanderthals and other non-modern Eurasians.

These pages are for the use of PNAS Editorial Board members and authors searching for PNAS member editors. For information about the National Academy of Sciences or its membership, please see
National Academy of Sciences | Copyright ©2022, All Rights Reserved